National parks and other large protected areas play an increasingly important role in the context of global social and environmental challenges. Nevertheless, they continue to be rooted in local places and cannot be separated out from their socio-cultural and historical context. Protected areas furthermore are increasingly understood to constitute critical sites of struggle whereby the very meanings of nature, landscape, and nature-society relations are up for debate. This paper examines governance arrangements and discursive practices pertaining to the management of the Danish Wadden Sea National Park and reflects on the relationship between pluralist institutional structures and pluralist, relational understandings of nature and landscape.